Identification of Cd36 (Fat) as an insulin-resistance gene causing defective fatty acid and glucose metabolism in hypertensive rats

Timothy J. Aitman, Anne M. Glazier, Caroline A. Wallace, Lisa D. Cooper, Penny J. Norsworthy, Faisal N. Wahid, Khulood M. Al-Majali, Paul M. Trembling, Christopher J. Mann, Carol C. Shoulders, Daniel Graf, Elizabeth St. Lezin, Theodore W. Kurtz, Vladimir Kren, Michal Pravenec, Azeddine Ibrahimi, Nada A. Abumrad, Lawrence W. Stanton, James Scott

Research output: Contribution to journalArticlepeer-review

652 Scopus citations

Abstract

The human insulin-resistance syndromes, type 2 diabetes, obesity, combined hyperlipidaemia and essential hypertension, are complex disorders whose genetic basis is unknown. The spontaneously hypertensive rat (SHR) is insulin resistant and a model of these human syndromes. Quantitative trait loci (QTLs) for SHR defects in glucose and fatty acid metabolism, hypertriglyceridaemia and hypertension map to a single locus on rat chromosome 4. Here we combine use of cDNA microarrays, congenic mapping and radiation hybrid (RH) mapping to identify a defective SHR gene, Cd36 (also known as Fat, as it encodes fatty acid translocase), at the peak of linkage to these QTLs. SHR Cd36 cDNA contains multiple sequence variants, caused by unequal genomic recombination of a duplicated ancestral gene. The encoded protein product is undetectable in SHR adipocyte plasma membrane. Transgenic mice overexpressing Cd36 have reduced blood lipids. We conclude that Cd36 deficiency underlies insulin resistance, defective fatty acid metabolism and hypertriglyceridaemia in SHR and may be important in the pathogenesis of human insulin-resistance syndromes.

Original languageEnglish
Pages (from-to)76-83
Number of pages8
JournalNature Genetics
Volume21
Issue number1
DOIs
StatePublished - Jan 1999

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